CHAPTER 28   THE REPRODUCTIVE SYSTEM

I. MALE REPRODUCTIVE SYSTEM

          A.Anatomy

                    1.testes (male gonads)

                              a.contained in the scotum(skin,fascia)

                                        (1)testes must descend for proper sperm production

                                                  (a)lower temperature below 37 C

                                        (2)muscle

                                                  (a)dortos

                                                            i)Sm M layer - wrinkles

                                                  (b)cremaster

                                                            i)Sk M - suspends testes

                                        (3)spermatic cord

                                                  (a)CT with nerves, vessels

                                                            i)BV: testicular arteries and veins

                              b.two tunics

                                        (1)tunica vaginalis - outer

                                        (2)tunica albuginea

                                                  (a)reach inward to divide into lobules

                                                            i)4 seminerferous tubules/lobules

                                                                      a)ST produce sperm

                                                  (b)interstitial cells (Leydig cells)

                                                            i)produce testerterone

                                                            ii)located in spaces between seminerferous tubules

                              c.rete testis

                                        (1)junction of seminerferous tubule lobules

                                        (2)sperm leave via efferent ductules into epididymis

                    2.epididymis

                              a.superior to testes

                                        (1)distal portion is posterolaterally located

                              b.immature sperm stored

                              c.mature on trip through the vas deferens (ductus)

                    3.vas deferens

                              a.runs superior through inguinal canal

                              b.enlarged region at distal end is ampulla

                              c.seminal vesicle joins to form ejaculatory duct

                              d.junction is at prostate gland

                                        (1)also join urethra from bladder (Prostatic urethra)

                              e.duct functions to transport and propel sperm to

                                        urethra (membranous to penile urethra)

                    4.seminal vesicles

                              a.2 glands

                              b.yellowish viscous alkaline fluid semen component

                              c.contains fructose

                    5.prostate gland

                              a.1 gland size of chestnut

                              b.milky alkaline fluid

                              c.enzymes to activate sperm

                              d.prostate problems effect urination

                    6.bulbourethral glands (Cowper's)

                              a.2 pea sized

                              b.thick, clear mucus

                              c.neutralize urethra before sperm pass

                    7.Penis

                              a.expernal genitalia

                              b.glans penis - enlarged tip

                              c.prepuce - foreskin

                                        (1)removed with circumcision

                              d.corpus spongiosum

                                        (1)flexible, erectile tissue

                                        (2)surrounds penile urethra

                              e.corpus cavernosa

                                        (1)erectile tissue

          B.Sperm formation

                    1.spermatogenesis

                              a.formation of sperm (gametes)

                              b.puberty

                    2.Meiosis

                              a.I and II to get 4 daughter cells instead of 2

                              b.haploid # chromosomes (23) n

                              c.normal cells diploid 2n (46) (23 pair of homologous chromosomes)

                              d.n from sperm + n from egg = 2n zygote

                              e.characteristics of meiosis I

                                        (1)PMAT

                                        (2)crossing over with homologous chrom. of tetrad in P

                                        (3)daughter cells not identical

                                                  (a)each has 1/2 of homologous chrom.pair

                                                  (b)n

                              f.characteristics of meiosis II

                                        (1)PMAT

                                        (2)daughters from I divide and replicated

                                                  pair before I finally separate

                                        (3)gives 4 daughter cells each with n chromosome

                    3.spermatogonia

                              a.stem cells undergo mitosis

                    4.spermatocytes (primary)

                              a.from meiosis I form haploid secondary spermatocytes

                              b.after meiosis II have spermatids

                              c.spermiogenesis

                                        (1)transformation to a tail

                              d.sperm head contains DNA

                              e.acrosome hydrolytic enzymes to penetrate egg

                    5.Sertoli cells

                              a.supporting cells in seminerferous tubules

                              b.nutrients supplied

                              c.barrier to prevent blood interaction

                                        (1)formation of antibodies

          C.Male sexual response

                    1.erection

                              a.vasodilation - blood flow increases

                              b.nitric oxide

                              c.corpora cavernosum and spongiosum

                              d.stimulation

                                        (1)mechanical

                                        (2)sensory

                                        (3)parasympathetic

                    2.ejaculation

                              a.propulsion of semen

                              b.sympathetic nerve outflow

                              c.contraction of ducts, glands, penile muscles

                              d.rapid HR, elevated BP

                              e.orgasm or climax

          D.Male Hormonal Regulation

                    1.Brain-Testicular axis

                              a.androgen (male sex hormones) production by

                                        (1)testes

                                        (2)hypothalamus

                                        (3) ant. pituitary

                    2.GnRH

                              a.hypothalamus

                              b.gonadotropin release from ant. pit

                                        (1)FSH

                                        (2)LH

                                        (3)enter circulation

                    3.FSH

                              a.stimulates spermatogenesis in testes along with testosterone

                    4.LH

                              a.sitmulates interstitial cells to secrete testosterone and small amount of estrogen

                              b.testosterone effects testes and enters circulation

                    5.Inhibin

                              a.feedback inhibition of hypothalamus and ant pituitary

                              b.produced by Sertoli cells in testes

                    6.Testerone

                              a.steroid hormone from cholesterol

                              b.induces gene expression to make proteins

                              c.can convert to

                                        (1)dihydrotestosterone in prostate cells

                                        (2)estrogen in brain neurons

                              d.initiates spermatogenesis at puberty

                              e.body anabolic effects

                              f.male sexual function

                              g.secondary sex characteristics

                                        (1)pubic,axillary,facial hair etc.

                                        (2)larynx enlarges

                                        (3)bones increase in density

                                        (4)stimulates Sk M growth

                                        (5)behavior

                                        (6)increases BMR

 

II.FEMALE REPRODUCTIVE SYSTEM

          A.ANATOMY

                    1.Ovaries

                              a.female gonads, produce gamete = ovum

                              b.produce estrogens and progesterone

                              c.held in place by

                                        (1)broad ligament

                                                  (a)suspensory ligament                                                                           

(2)mesovarium

                                        (3)ovarian ligament

                              d.tunica albuginea

                                        (1)external fibrous CT

                                        (2)covered by germinal epithelium

                              e.ovarian follicles

                                        (1)within cortex of ovary

                                        (2)each contains an oocyte,ovum

                                        (3)different stages identified as

                                                  (a)primordial follicle

                                                            i)l layer cells

                                                                      a)follicle cells

                                                  (b)primary follicle

                                                            i)2+ layers of cells 

                                                                      a)granulosa cells

                                                  (c)secondary follicle

                                                            i)fluid-filled spaces between granulosa cells

                                                            ii)central fluid filled space

                                                                      a)antrum

                                                  (d)Graafian follicle

                                                            i)oocyte sits on one side of antrum

                                                            surrounded by granulosa cells      

                                        (4)Corpus luteum

                                                  (a)follicle after ovulation

                                                  (b)degenerates

                    2.fallopian tubes (oviducts, uterine tubes)

                              a.each 4"

                              b.fertilization occurs

                              c.Fimbriae

                                        (1)finger-like projections over surface of ovary

                                        (2)widens to infundibulum to ampulla

                                        (3)narrows at isthmus and enters uterus

                              d.not continuous with ovary and increases risk of infection of

peritoneal cavity

                                                  (a)PID- microbes can enter

                                                            i)Pelvic inflammatory disease

                    3.Uterus

                              a.anterior to rectum

                              b.posterior to bladder

                              c.body

                              d.fundus

                              e.isthmus

                              f.cervix

                              g.wall

                                        (1)perimetrium        

                                                  (a)outermost serous layer

                                                  (b)visceral peritneum

                                        (2)myometrium

                                                  (a)SM M middle layer

                                        (3)endometrium

                                                  (a)mucosal lining

                                                  (b)simple columnar epithelium

                                                  (c)thick lamina propria

                                                  (d)implantation of embryo

                                                  (e)two layers

                                                            i)stratum functionalis

                                                                      a)sheds with menstruation

                                                            ii)stratum basalis

a)forms new functionalis after menstruation

                              h.supports

                                        (1)mesometrium

                                                  (a)part of broad ligament to support uterus laterally

                                        (2)uterosacral ligaments

                                                  (a)posteriorly

                                        (3)round ligaments

                                                  (a)anteriorly

                              i.pap smear

                                        (1)to detect abnormal cells

                    4.vagina

                              a.fibromuscular tube

                              b.wall

                                        (1)adventitia

                                                  (a)fibroelastic

                                        (2)muscularis

                                                  (a)SmM

                                        (3)mucosa

                              c.mucus from cervical mucosal glands

                              d.hymen

                                        (1)covering of orifice

                                        (2)ruptures

                    5.external genitalia

                              a.mons pubis

                                        (1)rounded area over pubic symphysis

                              b.labia majora

                              c.labia minora

                                        (1)enclose vestibule containing urethra opening and vaginal

orifice

                              d.greater vestibular (Bartholin'a glands )

                                        (1)like bulbourethral grands

                                        (2)mucus into vestibule

                              e.clitoris

                                        (1)erectile tissue

                                                  (a)corpora cavernosa

                              f.perineum

                                        (1)diamond region from anus to Mons pubis

                    6.Mammary glands

                              a.modified sweat glands

                              b.produce milk

                              c.areola

                                        (1)pigmented

                              d.nipple

                              e.internal during pregnancy, lactation

                                        (1)lobes

                                                  (a)lobules

                                                            i)alveoli

                                                                      a)secrete milk

                                                  (b)milk exits via lactiferous ducts to

                                                  (c)lactiferous ampulla

          B.Physiology of female reproductive system

                    1.oogenesis

                              a.# eggs determined at birth

                              b.oogonium - stem cell - diploid

                                        (1)mitosis before birth to produce

                                        (2)primary oocyte within follicle

                                        (3)grows and stops at birth

c.1 primary oocyte selected each month to enter meiosis I and forms Graafian follicle

                              d.divides into 2 daughters each haploid

                                        (1)secondary oocyte - ovulated

                                        (2)polar body - one not chosen

                              e.secondary oocyte sperm enters

                                        (1)Meiosis II initiated by fertilization

                                        (2)divides

                                                  (a)get 1 ovum (with sperm)

                                                  (b)1 polar body

                              f.polar body also divides to 2 more polar bodies

          Compare oogenesis with spermatogenesis.

                   

                    2.Hormones

                              a.GnRH - Hypothalmus

                              b.LH - ant. pituitary

(1)promote maturing of follicles to secondary                                   follicles in ovaries

                                        (2)high levels induce ovulation

                              c.FSH - ant. pituitary

                                        (1)stimulates developement of primary                                                                          follicles to secodary follicles

                              d.estrogen (E)

                                        (1)ovarian follicles secrete

                                        (2)growth of reproductive organs

                                        (3)regeneration of endometrium

                              e.progesterone (P)

                                        (1)major source is corpus luteum in ovaries

                                        (2)produced by placenta

                                        (3)regeneration of endometrium 

                                        (4)maintains uterus in pregnancy

                                        (5)prepares breasts for lactation

                              f.LH ,FSH, E high blood levels before ovulation

                                        (1)LH level large spike prior to ovulation

                              g.progesterone levels high following ovulation

                                        (1)preparing "for gestation"

                              h.fluctuation hormone levels have other roles

                    3.Ovarian Cycle (28 day cycle)

                              a.Follicular phase

                                        (1)maturing of follicle

                                        (2)zona pellucida membrane around follicle

                                        (3)corona radiata capsule of granulosa cells

                              b.ovulatory phase

                                        (1)release of oocyte

                                        (2)day 10-14

                                        (3)mittelschmerz with ovulation

                                        (4)one follicle is selected? over others that                                                                    are simultaneously maturing

(5)fraternal twins if 2 ovum released                                      and fertilized by 2 different sperm, same as siblings

                                        (6)identical twins

                                                  (a)1 ovum, 1 sperm

                              c.luteal phase

                                        (1)follicle after ovulation

                                        (2)corpus luteum forms\

                                                  (a)secretes progesterone,some estrogen

                                                  (b)degenerates to corpus albicans

                                        (3)14 day duration

                    4.Uterine Cycle

                              a.endometrial functional layer changes in nonpregnant

                              b.menstrual phase (sloughing off)

                              c.proliferative phase (mitosis,thickens)

                              d.secretory phase (P,E influence

                                        (1)occurs during luteal phase of cycle

          C.Female Sexual Response

                    1.rhythmic contraction of uterus

                    2.pleasure sensation and relaxation - orgasm

                    3.orgasm not necessary for conception

          D.Development

                    1.X, Y chromosomes

                              a.XX

                              b.XY

                    2.embryono differentiates sexually 6 week

                    3.Testes descend to scotum 2 months before birth

                              a.guided by gubernaculum

                    4.puberty

                              a.10-15 yrs

                              b.increasing levels of E and T

                              c.menses: menarche to menopause